The article is focused on grammatical load and phonological substratum of morphonological phenomena in the systems with dominant vowels (Greek, Gothic, German, and English). The synchronic analysis of these systems allows to oppose Greek to the Germanic languages on the following points: the vowel component is not the only component in the morphonological system, the morphonological phenomena expressing the Aspect and Voice (Greek); the vowel component is the only one in the morphonological system, the morphonological phenomena expressing the Tenses and Participles (Germanic languages). Diachronic analysis has shown possible ways of evolution: the loss of some morphonological oppositions due to the loss of the corresponding grammar categories (Gothic ? English); some morphonological oppositions drop out from the morphonological system (Ancient Greek ? Modern Greek); some new morphonological oppositions are included into the morphonological system (Gothic ? German).
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